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His television work includes the "Fire" episode of The X-Files, a year on the Jerry Bruckheimer action series Soldier of Fortune, guest-starring and recurring roles on The Practice, The Invisible Man, Special Unit 2, JAG, Star Trek: Voyager, The Chronicle, Monk, Las Vegas, CSI: NY, Chuck, CSI: Crime Scene Investigation and White Collar, among others. Mark played a demon named Arnon on an episode titled "Witches in Tights" on the TV series Charmed. He played Badger, a semi-comical cockney-style crime boss in the short-lived Joss Whedon show Firefly and later cast again in another Joss Whedon show, Dollhouse as Paul Ballard's dismissive FBI superior. He has appeared as a villain in season five of the Fox show 24 and as Patricia Arquette's serial-killer nemesis on Medium. He appeared as Romo Lampkin in seasons three and four of Battlestar Galactica, and had a recurring role as Manservant Neville in ABC Family's short-lived cult hit The Middleman. He has been seen as Anthony Anthros on Bionic Woman, as Sterling the nemesis character on Leverage, as the King of Hell Crowley on Supernatural, and as bank robber Tom Prescott on an episode of Burn Notice. He appeared in the pilot episode of the US series White Collar as a villainous master forger and in NBC's Chuck as the Director of the Ring criminal organisation. He appeared in the SyFy series Warehouse 13 as Regent Benedict Valda. He appears in the 2011 Doctor Who episodes "The Impossible Astronaut" and "Day of the Moon" playing the character Canton Everett Delaware III. This was the first time Sheppard had appeared in a British TV series. In October 2013, Sheppard returned to White Collar, reprising his 2009 role as master forger Curtis Hagen.